Koko the gorilla has died. Here's how she changed humanity for the better.

Koko was an incredible icon for the animal world. She will be missed.

The lowland gorilla who wowed scientists and the public alike with her mastery of sign language passed away on June 21 at age 46.

From the time of her birth, Koko was an instant animal celebrity. She was on the cover of National Geographic twice and became a symbol for those working to improve our understanding of animals and how we treat them.


In her later years, Koko stayed in the spotlight. As recently as 2016, she was making Instagram videos with the band The Red Hot Chili Peppers and even learning how to play the bass guitar with the band's musician Flea.

Image via FolsomNatural/Flickr.

"Her impact has been profound and what she has taught us about the emotional capacity of gorillas and their cognitive abilities will continue to shape the world," the Gorilla Foundation said in a statement.

Koko's groundbreaking communication skills created invaluable bridges in the relationship between humans and animals.

Koko was best known for learning sign language. Dr. Francine Patterson famously taught a young Koko some simple words and phrases that helped launch a larger program at Stanford University in 1974.

Koko eventually learned to understand an estimated 2,000 English words and learned to sign 1,000 of her own. Patterson stayed with Koko for her entire life, and their relationship was chronicled in a 2016 documentary.

Koko famously had relationships with other human celebrities like Robin Williams — something he called "a mind altering experience." Williams and Koko become close over the years, and after Williams' death, Koko became visibly emotional when she was given the news he had died.

In a viral video that surfaced around the time of his passing, Koko and Williams are seen playing games with each other, and she even recognizes his face on the cover of a VHS tape of one of his movies.

Another favorite celebrity of Koko's was the inimitable Mister Rogers, who she shared some lovely moments with.

And while Koko was in many ways "adopted" by our collective culture, she mimicked human behavior in her own ways, famously asking for a pet kitten for Christmas in 1984. Her caretakers gave her a stuffed animal, but she held out for the real thing. She finally got her pet kitten a year later. She hilariously signed "obnoxious cat" when it playfully bit her.

Her life is a reminder that how we care for and learn from our fellow creatures is an evolving process.

Koko was an animal icon, but she was also more than that. Her contributions to science, communication, and understanding of the animal kingdom has been profound.

She had equally lasting effect on average people as well, creating empathy and compassion for creatures that were often portrayed as threatening. Her legacy is part of a larger relationship between humans and nature that is gradually improving as we educate ourselves about the amazing world that surrounds us.

President Biden/Twitter, Yamiche Alcindor/Twitter

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Today, President Biden spoke to these issues in straightforward language before signing four executive actions that aim to:

- promote fair housing policies to redress historical racial discrimination in federal housing and lending

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Upworthy and P&G Good Everyday wanted to help keep kindness going strong, so they partnered up to create the Lead with Love Fund. The fund awards do-gooders in communities around the country with grants to help them continue on with their unique missions. Hundreds of nominations came pouring in and five winners were selected based on three criteria: the impact of action, uniqueness, and "Upworthy-ness" of their story.

Here's a look at the five winners:

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Gates Foundation

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